| Tribe2 |
Genus |
Species
of Burch3 |
| Planorbis |
||
| Gyraulus |
circumstriatus, crista4 , deflectus, hornensis, parvus | |
| Biomphalaria |
||
| Biomphalaria5 |
glabrata, havanensis, obstructa | |
| Helisoma |
||
| Helisoma 6 |
ammon, anceps, binnyi, campanulata, columbiense, corpulentum, duryi, eucosmium, magnificum, multivolvis, occidentale, pilsbryi, pseudotrivolvis, scalare, subcrenatum, tenue, trivolvis, truncatum | |
| Promenetus |
exacuous, umbilicatellus | |
| Menetus |
brogniartiana, dilatatus, opercularis, sampsoni | |
| Planorbula |
armigera, campestris | |
| Vorticifex 7 |
effusa, solida | |
| Carinifex 8 |
newberryi | |
| Drepanotrema |
||
| Drepanotrema |
aeruginosum, cimex, kermatoides | |
| Insertae sedis 9 |
||
| Amphigyra |
alabamensis | |
| Neoplanorbis |
carinatus, smithi, tantillus, umbilicatus | |
| Notes (1) Hubendick, B. (1955) Phylogeny in the Planorbidae. Trans. Zool. Soc. London 28: 453-542. See the FWGNA archives for the essay that accompanied the posting of this classification on 11Apr08. (2) Subsequent authors added the suffix "ini" to Hubendick's tribe names. The Planorbis tribe became "Planorbini," and so forth. (3) These are the species recognized by J. B. Burch (1989) in his "North American Freshwater Snails" (Malacological Publications, Hamburg, MI) with a couple addenda. (4) While admitting that it is "doubtful if there is sufficient reason to retain separate genera," Hubendick preserved the separation of crista in the genus Armiger. Armiger was formally subsumed under Gyraulus by Meier-Brook (1983, Malacologia 24: 1 - 113). (5) Hubendick wrote, "The African genera Biomphalaria and Afroplanorbis are undoubtedly congeneric. The genera Australorbis and Tropicorbis, which are mainly South American, do not differ in any essential way from the African forms. Taphius and Platytaphius are distinctly delimited neither from Tropicorbis nor Australorbis. It is most likely that all of these can be placed in one genus, Biomphalaria." (6) D. W. Tayler (1966, Malacologia 4: 1 - 172) raised the Baker subgenus Planorbella to the genus level and split all Helisoma except H. anceps into it. Burch followed. (7) Hubendick suggested Parapholyx (Hanna 1922) here. But Burch observed (his note #60) that Meek's (1870) Vorticifex, although originally a fossil taxon, would have priority. We agree. (8) Carinifex was subordinated under Helisoma by Burch (his note #51), following Henderson (1931) and Taylor (1966). (9) The Coosa River endemic genera Amphigyra and Neoplanorbis were not treated by Hubendick in 1955. Burch followed Walter (1971, AMU Ann. Repts 37: 47-51) recognizing a tribe "Neoplanorbinae." |