Disclaimers: (1)
This key isn't literally dichotomous. (2) It is intended for use with
the 41 species and subspecies of freshwater gastropods inhabiting Atlantic drainages of Georgia,
plus 2 unconfirmed (U).
The management will not
be responsible for any loss, damage, or injury resulting from its
application elsewhere.
1a) Operculum present. . . . . . Subclass Prosobranchia (2)
1b) Operculum absent . . . . . . Subclass Pulmonata (21)
2a) Operculum multispiral [photo] .
. . (3)
2b) Operculum concentric [photo]
. . . (4)
2c) Operculum paucispiral [photo] . . . (9)
3a) Adult shell bearing two carina . . . . Valvata bicarinata.
3b) Adult shell bearing three carina . . . Valvata
tricarinata.
4a) Penis arising from the mantle edge, labial palps whiplike, eggs
calcareous . . . Family Ampullariidae (5)
4b) Penis modified from a right tentacle, labial palps not
whiplike, females ovoviviparous . . . Family Viviparidae (6)
5a) Shell whorls gently sloping, sutures not indented or
incised . . . Pomacea
paludosa (U).
5b) Sutures strongly incised . . . Pomacea
maculata.
6a) Shell width approximately equal to, or sometimes greater than shell
height . . . Viviparus
intertextus.
6b) Shell narrower than high . . . . (7)
7a) Shell with color bands . . . . Viviparus
georgianus.
7b) Shell unbanded . . . . (8)
8a) Adult strikingly large, shells greater than 40 mm standard
length . . . Cipangopaludina japonica.
8b) Adults smaller . . . Campeloma
decisum.
9a) Adults over 10 mm shell length, both sexes present, oviparous . . .
. . . . Family Pleuroceridae (10)
9b) Adults over 10 mm shell length, all female, ovoviviparous brooders
. . . Melanoides tuberculata (U).
9b) Adults under 10 mm shell length . . . . . Family Hydrobiidae (13)
10a) Shell entirely uncostate . . . . Pleurocera
proxima.
10b) Shell at least slightly costate . . . . (11)
11a) Shell strongly costate throughout . . . Pleurocera
catenaria catenaria.
11b) Shell with faint costae, usually around apex only . . . . . (12)
12a) Shell slightly carinate . . . . Pleurocera
catenaria dislocata.
12b) Shell entirely without carination (vicinity of Hawkinsville) . . .
Pleurocera
floridensis timidus.
13a) Penis simple, with a single duct . . . (14)
13b) Penis with a single duct and a glandular terminal lobe . . . (15)
13c) Penis with a single duct and fringed with numerous papillae . . . Littoridinops
tenuipes.
13d) Penis with two ducts . . . (19)
14a) Adult shell never greater than 6 mm maximum dimension . .
. Somatogyrus
virginicus.
14b) Adult shell typically greater than 6 mm maximum dimension . . . Gillia altilis.
15a) Umbillicus imperforate, adults can be greater than 4.5 mm . . . Notogillia sathon.
15b) Shell smaller, umbillicus at least slightly open . . . (16)
16a) Penis with a small terminal lobe, more than two additional
glandular areas, and large penial filament . . . Spilochlamys
turgida.
16b) Penis with a small terminal lobe and not more than two additional
glandular areas . . . (17)
16c) Penis with a large terminal lobe and a short filament . . . (18)
17a) Ogeechee River . . . Marstonia
halcyon.
17b) Springs and spring-fed tributaries of the Ocmulgee River . . . Marstonia
agarhecta.
17c) Endemic to Rock Springs, a tributary of the Oconee River . . . Marstonia
gaddisorum.
18a) Ogeechee River . . . . Floridobia
Species A.
18b) Southern coastal . . . Floridobia
floridana.
19a) Operculum entirely paucispiral . . . . Amnicola limosa.
19b) Operculum initially multispiral, becoming paucispiral . . . (20)
20a) Shell substantially higher than wide . . . . Lyogyrus granum.
20b) Shell roughly as wide as high, sometimes wider . . . Lyogyrus latus.
21a) Shell dextral . . . . Family Lymnaeidae (22)
21b) Shell sinistral, not planispiral . . . . Family Physidae (24)
21c) Shell sinistral, planispiral . . . . Family Planorbidae (27)
21d) Shell patelliform . . . . Family Ancylidae (33)
22a) Aperture greater than 50% shell height . . . Lymnaea
columella.
22b) Aperture less than 50% shell height . . . (23)
23a) First lateral tooth of the radula tricuspid . . . . Lymnaea humilis.
23b) First lateral tooth of the radula bicuspid . . . . Lymnaea
cubensis/viator. [photos]
24a) One-part penial sheath . . . . . Physa acuta.
24b) Two-part penial sheath . . . . (25)
25a) Shell slender, body generally dark . . . . Physa carolinae.
25b) Animal lightly pigmented, Shell with shoulders more pronounced . .
. (26)
26a) Whorls convex . . . Physa
gyrina.
26b) Whorls not notably convex . . . Physa pomilia.
27a) Adult greater than 6 mm shell diameter . . . . (28)
27b) Adult less than 6 mm shell diameter . . . . (30)
28a) Shell easily crushed with fingers . . . Biomphalaria
havanensis.
28b) Shell stronger . . . . (29)
29a) Spire appears deeply indented when viewed from either aspect . . .
Helisoma
anceps.
29b) Spire appears flattened when viewed from one aspect . . . (30)
30a) Juvenile shell bearing threadlike striae and acute keel .
. . Helisoma
trivolvis.
30b) Juvenile shell glossy, no keel [figure]
. . . Helisoma scalare duryi.
31a) Shell with rounded periphery . . . Gyraulus parvus.
31b) Shell carinate . . . (32)
32a) Carina strong, approximately mid-whorl . . . Promenetus
exacuous.
32b) Carina weak, distinctly off mid-whorl . . . Menetus
dilatatus.
33a) Apex bearing fine radial striae . . . Ferrissia
fragilis.
33b) Apex entirely smooth . . . . (34)
34a) Apex approximately in the midline . . . . Laevapex fuscus.
34b) Apex distinctly to the right of midline . . . Hebetancylus
excentricus.