Disclaimers:
(1) This key isn’t literally dichotomous. (2) It is
intended for use with the 39 species and subspecies of freshwater
gastropods inhabiting the state of South
Carolina. The
management will not be responsible for any loss, damage, or injury
resulting from its application elsewhere.
1a) Operculum present. . . . . . Subclass Prosobranchia (2)
1b) Operculum absent . . . . . . Subclass Pulmonata (18)
2a) Operculum multispiral [photo] .
. . (3)
2b) Operculum concentric [photo]
. . . (4)
2c) Operculum paucispiral [photo] . . . (11)
3a) Adult shell bearing two carina . . . Valvata
bicarinata.
3b) Adult shell bearing three carina . . . Valvata
tricarinata.
4a) Penis arising from the mantle edge, labial palps whiplike, eggs
calcareous . . . Family Ampullariidae (5)
4b) Penis modified from a right tentacle, labial palps not
whiplike, females ovoviviparous . . . Family Viviparidae (6)
5a) Shell whorls gently sloping, sutures not indented or incised . . . Pomacea
paludosa.
5b) Sutures strongly incised . . . Pomacea maculata.
6a) Aperture rounded . . . (7)
6b) Aperture oblong, noticably longer than wide . . . (10)
7a) Adults strikingly large, shells in excess of 40 mm
standard length . . . Cipangopaludina
japonica.
7b) Adults smaller . . . Viviparus
(8)
8a) Shell width approximately equal to, or sometimes greater
than, shell height . . . Viviparus
intertextus.
8b) Shell narrower than high . . . (9)
9a) Shell rounded, with color bands . . . Viviparus
georgianus.
9b) Shell with flattened sides, unbanded . . . Viviparus subpurpureus.
10a) Shell bearing spiral cords . . . Lioplax
subcarinata.
10b) Shell smooth . . . Campeloma
decisum decisum.
11a) Adults over 10 mm shell length, all female, ovoviviparous
brooders. . .
Melanoides tuberculata.
11b) Adults over 10 mm shell length, both sexes present, oviparous . .
. . . Family Pleuroceridae (12)
11c) Adults under 10 mm shell length . . . . . Hydrobioid
families (13)
12a) Shell uncostate . . . . Pleurocera
proxima.
12b) Shell costate (sometimes faint, around apex only).
Spiral
cords, if present, restricted to juvenile whorls only . . . Pleurocera
catenaria dislocata.
12c) Shell costate, bearing spiral cords throughout . . . Pleurocera
catenaria catenaria.
13a) Penis simple . . . (14)
13b) Penis bifurcated . . . (15)
13c) Penis fringed with numerous papillae . . . (17)
14a) Adults never greater than 6 mm . . . Somatogyrus
virginicus.
14b) Adults generally greater than 6 mm . . . Gillia altilis.
15a) Operculum entirely paucispiral . . . . Amnicola limosa.
15b) Operculum initially multispiral, becoming paucispiral . . . (16)
16a) Shell length greater than shell width . . . Lyogyrus granum.
16b) Shell length equal to or greater than shell width . . . Lyogyrus
latus.
17a) Shells polymorphic, including morphs with carinae and
spines . . . Pyrgophorus parvulus.
17b) Shells never carinate or spiny . . .
Littoridinops
tenuipes.
18a) Shell dextral . . . . Family Lymnaeidae (19)
18b) Shell sinistral, not planispiral . . . . Family Physidae (21)
18c) Shell sinistral, planispiral . . . . Family Planorbidae (23)
18d) Shell patelliform . . . . Family Ancylidae (30)
19a) Aperture greater than 50% shell height . . . Lymnaea
columella.
19b) Aperture less than 50% shell height . . . (20)
20a) First lateral tooth of radula bicuspid . . . Lymnaea
cubensis/viator.
20b) First lateral tooth of radula tricuspid . . . Lymnaea humilis.
[photos]
21a) One part penial sheath . . . . . Physa acuta.
21b) Penial sheath constricted into two parts . . . (22)
22a) Shell slender, body generally dark . . . . Physa carolinae.
22b) Animal lightly pigmented, Shell with shoulders more pronounced . .
. Physa
pomilia.
23a) Adult greater than 5 mm shell diameter . . . . (24)
23b) Adult less than 5 mm shell diameter . . . . (28)
24a) Shell easily crushed with finger pressure . . . Biomphalaria
havanensis.
24b) Shell stronger . . . (25)
25a) Spire appears deeply indented when viewed from either aspect . . . Helisoma anceps.
25b) Spire appears flattened when viewed from one aspect . . . . (26)
26a) Juvenile shell bearing threadlike striae & acute
keel . . . Helisoma
trivolvis.
26b) Juvenile shell glossy, no keel [figure]
. . . (27)
27a) Shell higher than wide in aperture view . . . Helisoma scalare scalare.
27b) Shell wider than high in aperture view . . . Helisoma scalare duryi.
28a) Shell with rounded periphery . . . . Gyraulus parvus.
28b) Shell carinate . . . (29)
29a) Carina strong, approximately mid-whorl . . . Promenetus
exacuous.
29b) Carina weak, distinctly off mid-whorl . . . Menetus
dilatatus.
30a) Apex bearing fine radial striae . . . (31).
30b) Apex entirely smooth . . . . (32)
31a) Apex distinctly to the right of midline . . . Ferrissia
fragilis.
31b) Apex approximately in the midline . . . Ferrissia
rivularis. [photos]
32a) Apex approximately in the midline . . . . Laevapex fuscus.
32b) Apex distinctly to the right of midline . . . Hebetancylus
excentricus.