Disclaimers:
(1) This key isn't literally dichotomous. (2) It is intended for use
with the 38 species and subspecies of freshwater gastropods inhabiting Atlantic drainages
of the state of North Carolina, plus two species unconfirmed (U). The management will not be
responsible for any loss, damage, or injury resulting from its
application elsewhere.
1a) Operculum present. . . . . . Subclass Prosobranchia (2)
1b) Operculum absent . . . . . . Subclass Pulmonata (18)
2a) Operculum multispiral [photo] . . . Valvata
bicarinata.
2b) Operculum concentric [photo] . . . (3)
2c) Operculum paucispiral [photo] . . . (9)
3a) Penis
arising from the mantle edge, labial palps whiplike, eggs calcareous .
. . Pomacea maculata.
3b)
Penis modified from a right tentacle, labial palps not whiplike,
females ovoviviparous . . . Family Viviparidae (4).
4b) Shell narrower than high . . . . (5)
5a) Shell with color bands .
. . Viviparus
georgianus.
5b) Shell unbanded . . . . (6)
6a) Adult shell strikingly large, greater than 40 mm standard length .
. . (7)
6b) Adult shells smaller . . . (8)
7a) Shell uncarinate, aperture lip typically purple . . . Cipangopaludina
chinensis (U)
7b) Shell carinate, especially noticeable in juveniles, aperture never
purple . . . Cipangopaludina
japonica. [photo]
8a) Shell smooth . . . . Campeloma
decisum.
8b) Shell bearing fine spiral cords . . . . Lioplax
subcarinata.
9a) Adults over 10 mm shell length, all female, ovoviviparous brooders
. . . Melanoides tuberculata.
9b) Adults over 10 mm shell length, both sexes present, oviparous . . .
. . Family Pleuroceridae (10).
9c) Adults under 10 mm shell length . . . . . (14)
10a) Aperture height 50% of shell height or greater . . . . Leptoxis
carinata.
10b) Aperture height much less than 50% of shell height . . . . (11)
11a) Shell costate . . . . . . (12)
11b) Shell uncostate . . . . (13)
12a) Shell bearing spiral cords throughout . . . Pleurocera
catenaria catenaria.
12b) If spiral cords are present, they are restricted to juvenile
whorls only . . . Pleurocera
catenaria dislocata.
13a) Shell with one strong carination . . . . Pleurocera
proxima.
13b) Shell either entirely smooth, or with fine spiral cords . . . Pleurocera
virginica.
14a) Primarily terrestrial in life habit, found above water level . . .
Pomatiopsis
lapidaria (U)
14b) Entirely aquatic in life habit . . . Family Hydrobiidae (15)
15a) Penis simple, with a single duct . . . (16)
15b) Penis with a single duct and a glandular, terminal lobe . . . Floridobia
sp.
15c) Penis with a single duct and fringed with numerous papillae . . . Littoridinops
tenuipes
15d) Penis with two ducts . . . (17)
16a) Adults never greater than 6 mm . . . Somatogyrus
virginicus.
16b) Adults generally greater than 6 mm . . . Gillia altilis.
17a) Operculum entirely paucispiral . . . . Amnicola limosa.
17b) Operculum initially multispiral, becoming paucispiral . . . Lyogyrus granum.
18a) Shell dextral . . . . Family Lymnaeidae (19)
18b) Shell sinistral, not planispiral . . . . Family Physidae (21)
18c) Shell sinistral, planispiral . . . . Family Planorbidae (24)
18d) Shell patelliform . . . . Family Ancylidae (31)
19a) Aperture greater than 50% shell height . . . Lymnaea
columella.
19b) Aperture less than 50% shell height . . . (20)
20a) First lateral tooth of the radula tricuspid . . . . Lymnaea humilis.
20b) First lateral tooth of the radula bicuspid . . . . Lymnaea
cubensis/viator.
[photo]
21a) One-part penial sheath . . . Physa acuta.
21b) Two-part penial sheath . . . . (22)
22a) Shell slender, body generally dark. . . . Physa carolinae.
22b) Animal lightly pigmented, shell with shoulders more pronounced . .
. (23)
23a) Whorls convex . . . Physa
gyrina.
23b) Whorls not notably convex . . . Physa pomilia.
24a) Adult greater than 10 mm shell diameter . . . . (25)
24b) Adult less than 10 mm shell diameter . . . . (27)
25a) Spire appears deeply indented when viewed from either aspect . . .
Helisoma
anceps.
25b) Spire appears flattened when viewed from one aspect . . .
(26)
26a) Aperture strikingly flared . . . . Helisoma
magnifica.
26b) Aperture not flared . . . . Helisoma
trivolvis.
27a) Shell with rounded periphery . . . (28)
27b) Shell carinate . . . (30)
28a) Adults smaller than 4 mm maximum shell dimension . . . Gyraulus parvus.
28b) Adults larger than 4 mm maximum shell dimension . . . . (29)
29a) Whorls tightly concentric . . . Planorbula
armigera.
29b) Whorls expanding markedly with growth . . . Helisoma
eucosmium.
30a) Carina strong, approximately mid-whorl . . . Promenetus
exacuous.
30b) Carina weak, distinctly off mid-whorl . . . Menetus
dilatatus.
31a) Apex bearing fine radial striae . . . (32)
31b) Apex entirely smooth . . . . (33)
32a) Apex distinctly to the right of midline . . . Ferrissia
fragilis.
32b) Apex approximately in the midline . . . Ferrissia
rivularis.
33a) Apex approximately in the midline . . . . Laevapex fuscus.
33b) Apex distinctly to the right of the midline . . . Hebetancylus
excentricus.